Credit: Park et al.
Stratigraphically calibrated phylogeny of Sphenisciformes correlated with tectonic movements and changing ocean circulation in the southern hemisphere showing how: (1) the Australian taxa are dispersed across the phylogeny temporally; (2) the Australian continent becomes progressively more isolated from other southern continents; and (3) a strengthened ACC (indicated by the black arrows) provides a new dispersal vector to Australia despite the presence of a strengthening Antarctic Polar Front (APF).
The bottom palaeomaps are based on reconstructions in Lawver & Gahagan [9]. Penguin silhouettes show overall trend for decreasing body size in penguin evolution: Top, archaic giant stem penguin taxa; middle medium-sized stem penguin taxa; bottom, smaller crown penguin taxa (silhouette credit: Fir0002/Flagstaffotos (original photo), John E. McCormack, Michael G. Harvey, Brant C. Faircloth, Nicholas G. Crawford, Travis C. Glenn, Robb T. Brumfield & T. Michael Keesey, used under a CC BY 3.0 Attribution Unported Licence (http://creativecommons.org/licenses/by/3.0/))).
Palaeoceanographic reconstructions after [9,72-74]. Palaeoceanographic abbreviations: EAC = East Australian Current, pEAC = palaeo-East Australian Current, pRG = palaeo-Ross Sea Gyre/Tasman Current, RG = Ross Sea Gyre. The relative strength of the ACC and APF is shown by thickening arrows and lines though time. Black arrow = cold currents, red arrows = warm currents.